TOSAENSE Makino, 1901
Subgenus Sedum
Distribution : Japan (Shikoku), China (Zhejiang), on limestone rocks, rare.
Description (by 't Hart & Bleij in IHSP, 2003) :
Perennial loosely tufted herbs with ascending to decumbent slender fleshy stems, ± 12 cm tall.
Leaves alternate, spatulate, rounded or retuse at the tip, petiolate, with delicate veins.
Flowering branches numerous, ascending.
Inflorescences cymes with 1 - 3 short spreading 1- to 5-flowered cincinni, these sometimes forked, bracts linear-spatulate, obtuse, emarginate.
Flowers 5- merous, (sub-) sessile, sepals basally free, spurred, unequal, oblong-linear to linear-spatulate, obtuse, petals slightly connate at the base, oblong-lanceolate, acute, yellow, nectar scales minute, rectangular-spatulate, rounded to truncate, flat, styles short, ± curved, fruit spreading, seeds probably reticulate-papillose.
Close to Sedum tricarpum and regarded as a rare Japanese endemic by Ohba (2001: 22). The Japanese descriptions are all incomplete in regard of the characters of sepals and seeds, but Fu & Rao (1988: 121) noted that the Chinese specimens have spurred sepals (basally free) and that the seeds are densely mamillate (probably reticulate-papillose). The differences are not very clear, however, and may relate to the phyllotaxis.
Two subspecies :